The wild-type A produces the agouti shift phenomenon, which causes hairs to be banded with black and an orangish/reddish brown, this revealing the underlying tabby pattern (which is determined by the T alleles at the separate tabby gene). The Agouti gene, with its dominant A allele and recessive a allele, controls the coding for agouti signaling protein (ASIP Q865F0). This is probably related to the phenomenon known as "tarnishing" in silvers.Īgouti An Abyssinian kitten displaying the ticked tabby pattern These cats resemble shaded or tipped goldens, but are genetically shaded or tipped silvers. A modifying factor has also been hypothesized in shaded silver and chinchilla Persians whose fur turns pale golden in adulthood, due to low levels of phaeomelanin production. Like amber cats, russet cats lighten as they age. Another recessive mutation at extension was discovered which causes the russet color in Burmese cats.This phenomenon was first identified in Norwegian Forest cats. Paws and nose still exhibit the original undiluted color, this in contrast to other diluted colors, where paws and nose have the diluted color. Kittens are born dark but lighten up as they age. A mutation at the extension locus E/e (the melanocortin 1 receptor, MC1R) changes black pigment to amber or light amber, similar in appearance to red and cream.The existence of this phenomenon as a discrete gene is a controversial subject among feline enthusiasts. The Dilution modifier gene, Dm, "caramelizes" the dilute colors as a dominant trait. It is different from the browning gene and has only been observed in laboratory cats.
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